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Plant Pest Diagnostics Center

Shaun Winterton

Dr. Shaun L. Winterton

Senior Insect Biosystematist

Plant Pest Diagnostics Branch, California Department of Food & Agriculture
3294 Meadowview Road, Sacramento, CA 95832-1448, U.S.A.
916-262-1166 office, 916-262-1190 fax,

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Curriculum vitae


  • PhD, Insect Systematics & Evolution - Department of Zoology & Entomology, University of Queensland, Brisbane, Australia (2000)
  • Advanced Diploma of Aquatic Resource Management - Central Queensland University, Rockhampton, Australia (1999)
  • Post-Graduate Diploma in Science (Entomology, Systematics) - University of Queensland, Brisbane, Australia (1997)
  • Bachelor of Applied Science- (Zoology, Chemistry) - University of Southern Queensland, Toowoomba, Australia (1992)

Refereed Publications (reverse chronological)

  1. Zhao, J., Winterton, S.L. & Liu, Z. (2013) Ancestral gene organization in the mitochondrial genome of Thyridosmylus langii (McLachlan, 1870) (Neuroptera: Osmylidae) and implications for lacewing evolution. PLoS ONE, 8 (5): e62943 [doi:10.1371/journal.pone.0062943].
  2. Wang, Y., Liu, X., Winterton, S.L., Yan, Y., Chang, W., Yang, D. (2013) Comparative mitogenomic analysis reveals sexual dimorphism in a rare montane lacewing (Insecta: Neuroptera: Ithonidae). PLoS One submitted.
  3. Winterton, S.L. (2013) A new species of Agapophytus Guérin named for the late Donald Colless (Diptera: Therevidae: Agapophytinae). Zootaxa, 3680, 190-194 [doi:10.11646/zootaxa.3680.1.12]
  4. Sziráki, G. & Winterton, S.L. (2012) An unusual Cryptoscenea Enderlein (Neuroptera: Coniopterygidae) species from Western Australia. Genus, 23, 419–424. [PDF]
  5. Wang, Y., Liu, X., Winterton, S.L. & Yang, D. (2012) The first mitochondrial genome for the fishfly subfamily Chauliodinae and implications for the higher phylogeny of Megaloptera. PLoS ONE, 7(10): e47302. [doi:10.1371/journal.pone.0047302].
  6. Yang, Q., Makarkin, V.N., Winterton, S.L., Khramov, A.V. & Ren, D. (2012) A remarkable new family of Jurassic Insects (Neuroptera) with primitive wing venation and its phylogenetic position in Neuropterida. PLoS ONE 7(9): e44762. [doi:10.1371/journal.pone.0044762]
  7. Winterton, S.L., Guek, H.P. & Brooks, S.J. (2012) A charismatic new species of green lacewing discovered in Malaysia (Neuroptera: Chrysopidae): the confluence of citizen scientist, online image database and cybertaxonomy, Zookeys, 214, 1–11. [doi: 10.3897/zookeys.214.3220]
  8. Winterton, S.L. (2012) Review of Australasian spider flies (Diptera: Acroceridae) with a revision of Panops Lamarck. Zookeys, 172, 7–75. [doi: 10.3897/zookeys.172.1889]
  9. Winterton, S.L. & Gillung, J.P. (2012) A new species of spider fly in the genus Sabroskya Schlinger from Malawi, with a key to Acrocerinae world genera (Diptera: Acroceridae). Zookeys, 171, 1–15. [doi: 10.3897/zookeys.171.2137]
  10. Winterton, S.L. & Lambkin, C.L. (2012) New Australian stiletto flies: revision of Manestella Metz and description of Medomega gen. nov. (Diptera: Therevidae: Agapophytinae). Zookeys, 240, 1-119. [doi: 10.3897/zookeys.240.2967]
  11. Winterton, S.L., Hauser, M. & Badrawy, H.B.M. (2012) A remarkable new genus of stiletto flies from Egypt, with a key to Palaearctic genera of Phycinae (Diptera: Therevidae). Zookeys, 184, 35–45. [doi: 10.3897/zookeys.184.2759]
  12. Winterton S.L. & Ferguson, D. (2012) New species of Vomerina Winterton (Diptera: Therevidae: Agapophytinae) from Australia. Zookeys 218, 65–75. [doi: 10.3897/zookeys.218.2380]
  13. Winterton, S.L. (2011a) Revision of the stiletto fly genera Acupalpa Kröber and Pipinnipons Winterton (Diptera, Therevidae, Agapophytinae) using cybertaxonomic methods, with a key to Australasian genera. Zookeys, 95, 29–78. [doi:10.3897/zookeys.95.1461]
  14. Winterton, S.L. (2011b) Review of the stiletto fly genus Actenomeros Winterton & Irwin (Diptera: Therevidae: Agapophytinae). Zookeys, 120, 55–63. [doi:10.3897/zookeys.120.1615]
  15. Winterton S.L. (2011c) New species of Prepseudatrichia Kelsey from Thailand (Diptera: Scenopinidae). Zookeys, 122, 39–44. [doi:10.3897/zookeys.122.1598]
  16. Winterton, S.L. & Gharali B. (2011) Iranotrichia gen. nov.: a new genus of Scenopinidae (Diptera) from Iran, with a key to window fly genera of the world. Zookeys, 138, 75–92. [doi:10.3897/zookeys.138.1821]
  17. Winterton, S.L. & Gaimari, S.D. (2011) Revision of South American window fly genus Heteromphrale Kröber, 1937 (Diptera: Scenopinidae). Zookeys, 84, 39–57. [doi:10.3897/zookeys.84.774]
  18. Gillung, J.P. & Winterton, S.L. (2011) New genera of philopotine spider flies (Diptera: Acroceridae) with a key to living and fossil genera. Zookeys, 127, 15–27. [doi:10.3897/zookeys.127.1824]
  19. Wang, Y, Winterton, S.L. & Liu, Z. (2011) Phylogeny and biogeography of Thyridosmylus (Neuroptera: Osmylidae). Systematic Entomology, 36, 330–339. [doi:10.1111/j.1365-3113.2010.00565.x]
  20. Winterton S.L. & Makarkin, V. (2010) Phylogeny of moth lacewings and giant lacewings (Neuroptera: Ithonidae, Polystoechotidae) using DNA sequence data, morphology and fossils. Annals of the Entomological Society of America, 103, 511–522. [doi:10.1603/AN10026]
  21. Winterton S.L., Hardy, N.B. & Wiegmann B.M. (2010) On wings of lace: phylogeny and Bayesian divergence time estimates of Neuropterida (Insecta) based on morphological and molecular data. Systematic Entomology, 35, 349–378. [doi:10.111/j.1365-3113.2010.00521.x]
  22. Winterton, S.L. (2010) A new species of Stenobiella Tillyard (Neuroptera: Berothidae) from Australia. Zookeys, 64, 1–8. [doi:10.3897/zookeys.84.774]
  23. Aldrich, J.R., Lee, T.C., Zhang, Q.H., Torres, J., Winterton, S.L., Han, B., Miller, G.L. (2009) Prothoracic gland semiochemicals of green lacewings (Neuroptera: Chrysopidae). Journal of Chemical Ecology, 35, 1181–1187. [doi:10.1007/s10886-009-9701-x]
  24. Winterton S.L. & Wiegmann B.M. (2009) Neuroptera. In S. Blair Hedges & Sudhir Kumar (Eds), The Time-tree of Life. Oxford University Press, New York. 572 pp.
  25. Winterton, S.L. (2009a) Revision of the stiletto fly genus Neodialineura Mann (Diptera: Therevidae: Agapophytinae): an empirical example of cybertaxonomy. Zootaxa, 2157: 1–33. [PDF]
  26. Winterton S.L. (2009b) ‘Scales and setae’. pp. 901-904. In R.T. Cardé & V.H. Resh (eds.), Encyclopedia of Insects. Academic Press/Elsevier Science, San Diego, USA. Encyclopedia of insects 2nd edition. 1024 pp.
  27. Winterton, S.L. (2009c) A new species of Pseudatrichia Osten Sacken (Diptera: Scenopinidae) from North America. Zootaxa, 2094: 36–41. [PDF]
  28. Winterton S.L. (2009d) Ch 14. Diptera Biodiversity Informatics. pp. 381–406. In T. Pape, D. Bickel & R. Meier (eds.). Diptera Diversity: Status, Challenges and Tools. Koninklijke Brill NV. pp. 449.
  29. Winterton, S.L. & Woodley N. (2009) New species of Metatrichia Coquillett (Diptera: Scenopinidae) from Australia and Venezuela. Zootaxa, 2094: 42–51. [PDF]
  30. La Salle, J., Wheeler, Q., Jackway, P., Winterton S., Hobern, D. & Lovell, D. (2009) Accelerating taxonomic discovery through automated character extraction. Zootaxa, 2217: 43–55. [PDF]
  31. Wiegmann, B.M., Trautwein, M.D., Kim, J., Cassel, B.K., Bertone, M.A., Winterton, S.L. & Yeates, D.K. (2009) Single-copy nuclear genes resolve the phylogeny of the holometabolous insect orders. BMC Biology, 7: 34. [doi:10.1186/1741-7007-7-34]
  32. Winterton, S.L. & Irwin, M.E. (2008) Kaurimyia gen. nov.: discovery of Apsilocephalidae (Diptera: Therevoid clade) in New Zealand. Zootaxa, 1779: 38–44. [PDF]
  33. Winterton, S.L. (2008) An unusual new species of Scenopinus Latreille (Diptera: Scenopinidae) from Australia. Zootaxa, 1895: 66–68. [PDF]
  34. Kerr, P.H. & Winterton, S.L. (2008) Do parasitic flies attack mites? Evidence in Baltic amber. Biological Journal of the Linnaean Society, 93: 9–13. [doi:10.1111/j.1095-8312.2007.00935.x]
  35. Winterton, S.L. (2007a) Revision of the Australian stiletto fly genus Acatopygia Kröber (Diptera: Therevidae: Agapophytinae). Zootaxa, 1405: 51–62. [PDF]
  36. Winterton, S.L. (2007b) New species of Laxotela Winterton & Irwin from Australia (Diptera: Therevidae: Agapophytinae). Zootaxa, 1407: 43–55. [PDF]
  37. Winterton, S.L. (2007c) New species of Nanexila Winterton & Irwin and Taenogera Kröber from Australia (Diptera: Therevidae). Zootaxa, 1413: 55–64. [PDF]
  38. Winterton, S.L. (2007d) New species of Acraspisoides Hill & Winterton and Bonjeania Irwin & Lyneborg (Diptera: Therevidae: Agapophytinae), with the description of a new genus. Zootaxa, 1438: 1–25. [PDF]
  39. Winterton, S.L., Wiegmann, B.M. & Schlinger, E.I. (2007) Molecular phylogeny and Bayesian divergence time estimates of small-headed flies (Diptera: Acroceridae). Molecular Phylogenetics & Evolution, 43, 808–832. [doi:10.1016/j.ympev.2006.08.015]
  40. Hauser, M. & Winterton, S.L. (2007) A new fossil genus of small headed fly (Diptera: Acroceridae) from Baltic amber. Annals of the Entomological Society of America, 100: 152–156. [doi: 10.1603/0013-8746(2007)100[152:ANFGOS]2.0.CO;2]
  41. Walter D.E. & Winterton S.L. (2007) Keys and the Crisis in Taxonomy: Extinction or Reinvention? Annual Review of Entomology, 52: 193–208. [doi:10.1146/annurev.ento.51.110104.151054]
  42. Penny, N.D. & Winterton S.L. (2007) Rediscovery of the unusual genus Ormiscocerus (Neuroptera: Berothidae: Cyrenoberothinae). Proceedings of the California Academy of Sciences, 58: 1–6. [PDF]
  43. Winterton, S.L. (2006a) New species of Eupsilocephala Kröber from Australia (Diptera: Therevidae). Zootaxa, 1372: 17–25. [PDF]
  44. Winterton, S.L. (2006b) Aberrant wing venation in Apochrysa lutea (Walker) (Neuroptera: Chrysopidae: Apochrysinae) from Australia. Australian Entomologist, 33: 143–146.
  45. Winterton, S.L. & DeFreitas S. (2006) Molecular phylogeny of the green lacewings (Neuroptera: Chrysopidae) Australian Journal of Entomology, 45: 235–243. [doi:10.1111/j.1440-6055.2006.00537.x]
  46. Winterton, S.L. & Kerr, P.K. (2006) A new species of Alloxytropus Bezzi (Diptera: Scenopinidae: Proratinae) from Israel. Zootaxa, 1155: 41–50. [PDF]
  47. Winterton, S.L. & Metz, M.A. (2005) Cyrtosathe gen. n.: the first non-scenopinine window-fly from sub-Saharan Africa (Diptera: Scenopinidae). Zootaxa 975: 1–12. [PDF]
  48. Winterton, S.L. (2005) A new species of Propebrevitrichia Kelsey (Diptera: Scenopinidae: Scenopininae) from Botswana. Zootaxa 818: 1–8. [PDF]
  49. Winterton, S.L. (2004a) ‘Lacewings (Neuroptera)’ In ‘Gzimek’s Animal Encyclopedia’ vol . 3. ‘Insects’ pp. 305– 314. Schlager Press, Chicago, USA.
  50. Winterton, S.L. (2004b) Are hind coxal knobs a synapomorphy for therevids?: An unusual new species of Anabarhynchus Macquart from Australia (Diptera: Therevidae: Therevinae).  Zootaxa 413: 1–8. [PDF]
  51. Hill, H.N. & Winterton, S.L. (2004) Acraspisoides- a new genus of Stiletto-flies from Australia (Diptera: Therevidae). Zootaxa 414: 1–15. [PDF]
  52. Winterton, S.L. (2003a) Molecular Phylogeny of Neuropterida with Emphasis on the Lacewings (Neuroptera). Entomologische Abhandlungen 61 (2): 158–160.
  53. Winterton, S.L. (2003b) ‘Scales and setae’. In R.T. Cardé & V.H. Resh (eds.), Encyclopedia of Insects. pp.1017-1020. Academic Press/Elsevier Science, San Diego, USA.
  54. Winterton, S.L. & Brooks, S.J. (2002) Phylogeny of the apochrysine green lacewings (Neuroptera: Chrysopidae: Apochrysinae). Annals of the Entomological Society of America 95: 16–28. [doi:10.1603/0013-8746(2002)095[0016:POTAGL]2.0.CO;2]
  55. Winterton, S.L. (2002a) ‘Molecular Systematics of Diptera’. In J.H. Skevington and P.T. Dang (eds), ‘Exploring the diversity of flies (Diptera)’. Biodiversity [feature article] 3(4): 21–22.
  56. Winterton, S.L. (2002b) ‘Aquatic plant cultivation’. In Weed Biocontrol: The theory and practical application of biological control of weeds. M. Julien & G. White (eds.). ACIAR Monograph.
  57. Lambkin, C. L., Lee, M.S., Winterton, S.L. & Yeates D.K. (2002) Partition Bremer Support and multiple trees. Cladistics 18: 436–444. [doi:10.1111/j.1096-0031.2002.tb00159.x]
  58. Metz, M.A., Winterton, S.L. & Irwin, M.E. (2002) Notes on the functional morphology of terminalia from Prorates ballmeri Nagatomi & Liu (Diptera: Scenopinidae: Proratinae) collected while in copula, with a description of the previously unknown female. Zootaxa 76: 1–12. [PDF]
  59. Winterton, S.L. & Irwin, M.E. (2001) Phylogenetic revision of Agapophytus Guérin (Diptera: Therevidae: Agapophytinae). Invertebrate Taxonomy 15: 467–526. [doi:10.1071/IT00027]
  60. Winterton, S.L., Yang, L., Wiegmann, B.M. & Yeates, D.K. (2001) Phylogenetic revision of Agapophytinae subf. n. (Diptera: Therevidae) based on molecular and morphological evidence. Systematic Entomology 26: 173–211. [doi:10.1046/j.1365-3113.2001.00142.x]
  61. Winterton, S.L. (2000) Phylogenetic revision of Acupalpa Kröber (Diptera: Therevidae: Agapophytinae). Insect Systematics and Evolution 31: 225–240. [doi: 10.1163/187631200X00417]
  62. Winterton, S.L., Skevington, J.H., Irwin, M.E. & Yeates, D.K. (2000) Phylogenetic revision of Bonjeania Irwin & Lyneborg(Diptera: Therevidae). Systematic Entomology 25: 295–324. [doi:10.1046/j.1365-3113.2000.00107.x]
  63. Heard, T.A. & Winterton, S.L. (2000) Interactions between nutrient status and weevil herbivory in the biological control of water hyacinth. Journal of Applied Ecology 37: 117–127. [doi: 10.1046/j.1365-2664.2000.00480.x]
  64. Winterton, S.L. (1999) Obligatory ontogenetic colour change correlated with sexual maturity in adult Chrysoperla congrua (Walker) (Neuroptera: Chrysopidae). Australian Journal of Entomology 38: 120–123. [doi:10.1046/j.1440-6055.1999.00090.x]
  65. Winterton, S.L. & Irwin, M.E. (1999) Laxotela- a new genus of Therevidae (Diptera) from Australia. Entomologica Scandinavica 30: 299–310. [doi:10.1163/187631200X00138]
  66. Winterton, S.L., Irwin, M.E. & Yeates, D.K. (1999a) Systematics of Nanexila gen. nov. (Diptera: Therevidae) from Australia. Invertebrate Taxonomy 13: 237–308. [doi:10.1071/IT97029]
  67. Winterton, S.L., Irwin, M.E. & Yeates, D.K. (1999b) Phylogenetic revision of the Taenogera Kröber genus-group (Diptera: Therevidae), with descriptions of two new genera. Australian Journal of Entomology 38: 274–290. [doi:10.1046/j.1440-6055.1999.00126.x]
  68. Winterton, S.L., Merritt, D., O’Toole, A., Irwin, M.E. & Yeates, D.K. (1999c) Morphology and histology of the spermathecal sac, a novel structure in the female reproductive system of Therevidae (Diptera: Asiloidea). International Journal of Insect Morphology and Embryology 28: 273–279. [doi:10.1016/S0020-7322(99)00030-6]
  69. Winterton, S.L. (1997) ‘Aquatic plant cultivation’. In M. Julien & G. White (eds), Biological Control of Weeds: theory and practical application. pp.: 151–156. ACIAR Monograph, 49.
  70. Winterton, S.L. & Brancatini,V.A. (1996) New information on Brinckochrysa Tjeder and Ankylopteryx Brauer (Neuroptera: Chrysopidae), with descriptions of genitalia of A. basalis Kimmins and A. rieki New. Australian Entomologist 23: 21–26.
  71. Winterton, S.L. (1995a) A new species of Mallada Navás (Neuroptera: Chrysopidae) from Australia with a key to species. Journal of the Australian Entomological Society 34: 23–27. [doi:10.1111/j.1440-6055.1995.tb01270.x]
  72. Winterton, S.L. (1995b) A new genus and species of Apochrysinae (Neuroptera: Chrysopidae) from Australia, with a checklist of Australian Chrysopidae. Journal of the Australian Entomological Society 34: 139–145. [doi:10.1111/j.1440-6055.1995.tb01306.x]
  73. Winterton, S.L. (1995c) Sexual Dimorphism in Mallada traviata (Banks) (Neuroptera: Chrysopidae). Australian Entomologist 22:123–124.
  74. Lambkin, K.J. & Winterton, S.L. (1995) Chapter 26, ‘Lacewings (Neuroptera)’. In Wildlife of Greater Brisbane. Queensland Museum Publication.
  75. Murray, D.A.H., Rynne, K.P., Winterton, S.L., Bean, J.A. & Lloyd, R.J. (1995) Effect of host plant on parasitism of Helicoverpa armigera (Hübner) (Lepidoptera: Noctuidae) by Hyposoter didymator Thunberg (Hymenoptera: Ichneumonidae) and Cotesia kazak (Telenga) (Hymenoptera: Braconidae). Journal of the Australian Entomological Society 34: 71–73. [doi:10.1111/j.1440-6055.1995.tb01284.x]
  76. Winterton, S.L. (1994) Cryptocoryne Fischer: aspects of aquatic plant growth and ecology. Superfish (Queensland Cichlid Group publication) Sept-October issue, pp.: 6–9.

Electronic Publications (Chronological)

  1. Winterton, S.L. (2004) ‘Aquarium and Pond Plants of the World’. Lucid key. Animal and Plant Health Inspection Service (APHIS), United States Department of Agriculture (USDA), ver 1. (CD publication).
  2. Winterton, S.L., Skevington, J.H. & Lambkin, C.L. (2005) ‘Stiletto flies of Australasia (Diptera: Therevidae)’. Online Lucid3 Interactive key and website.  California Department of Food & Agriculture, Agriculture Canada and CSIRO. ver 1. [Link]
  3. Winterton, S.L. & J. Scher (2006) ‘Aquarium and Pond Plants of the World. 2nd edition’. Lucid key. Animal and Plant Health Inspection Service (APHIS), United States Department of Agriculture (USDA), ver 1. (CD publication). Revised and upgraded version of the popular interactive key to world genera of aquatic plants traded in the aquarium and pond plant industry for quarantine officials. [Link]