Bonjeania Irwin & Lyneborg

Lonchorhynchus White, 1915: 33 (preoccupied Wiman, 1910); replacement name- Bonjeania Irwin & Lyneborg, 1989: 356. Type species: Anabarhynchus nitidifrons Macquart, 1850: 403.

Could be confused with: Parapsilocephala, Patanothrix, Acatopygia, Laxotela.

Systematic grouping: Agapophytinae.

Diagnosis: Antenna positioned low on frons; antenna shorter than head length; male occiput concave with single row of postocular setae; female occiput convex with multiple, poorly defined rows of postocular setae; wing cell m3 closed; wing infuscation usually sexually dimorphic; anteroventral apical seta absent, fore and hind femora with velutum patches; posterior margin of abdominal tergites 1-4 often with thickened band of velutum orientated anteriorly; medial atrium present; gonocoxite with velutum (often on membrane bordering medial atrium); gonocoxal apodemes usually greatly elongate, greater than 4 times length of gonocoxite, rarely shorter; distiphallus usually extremely long and thin; ejaculatory apodeme and lateral ejaculatory apodeme greatly enlarged; single spermatheca; spermathecal sac simple, spherical (rarely reduced).

Distribution: Australia.

Comments: Bonjeania is the sister-genus to Parapsilocephala, but has highly modified male genitalia and female spermathecal sac. There are at least six undescribed species represented in collections. The genus is presently being revised by SLW. Spot characters to look for when identifying this genus include velutum patches on the fore and hind femora, wing cell m3 closed, single spermatheca and often greatly enlarged gonocoxal and ejaculatory apodemes.

Included species: B. actuosa (White), B. angelikae Winterton & Skevington, B. clamosis Winterton & Skevington, B. dynastis Winterton & Skevington, B. flavofemoralis Winterton & Skevington, B. irwinae Winterton & Skevington, B. nitidifrons (Macquart), B. orphne Winterton & Skevington, B. segnis (White), B. trilineata Winterton & Skevington.

Relevant literature:
Macquart, J.M. (1850) Diptères exotiques nouveaux ou peu connus. 4.e supplement. Mémoirs de la Société Royale des Sciences, de l’Agriculture et des Arts, de Lille, 1849, 309-479.

Mann, J.S. (1929) Revisional notes on Australian Therevidae. Part 2. Australian Zoologist, 6, 17-49.

White, A. (1915) The Diptera-Brachycera of Tasmania. Part II. Families Tabanidae and Therevidae. Papers and Proceedings of the Royal Society of Tasmania, 1915, 2-60.

Winterton, S.L., Skevington, J.H., Irwin, M.E. & Yeates, D.K. (2000) Phylogenetic revision of Bonjeania Irwin & Lyneborg (Diptera: Therevidae). Systematic Entomology 25: 295-324.